the perpetuation of living beings-第4章
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which is largely marked with it。 In this case; as I have said; there
was no means of the second generation intermarrying with any but
five…fingered persons; and the question naturally suggests itself; What
would have been the result of such marriage? Reaumur narrates this
case only as far as the third generation。 Certainly it would have been
an exceedingly curious thing if we could have traced this matter any
further; had the cousins intermarried; a six…fingered variety of the
human race might have been set up。
To show you that this supposition is by no means an unreasonable one;
let me now point out what took place in the case of Seth Wright's
sheep; where it happened to be a matter of moment to him to obtain a
breed or raise a flock of sheep like that accidental variety that I
have describedand I will tell you why。 In that part of Massachusetts
where Seth Wright was living; the fields were separated by fences; and
the sheep; which were very active and robust; would roam abroad; and
without much difficulty jump over these fences into other people's
farms。 As a matter of course; this exuberant activity on the part of
the sheep constantly gave rise to all sorts of quarrels; bickerings;
and contentions among the farmers of the neighbourhood; so it occurred
to Seth Wright; who was; like his successors; more or less 'cute; that
if he could get a stock of sheep like those with the bandy legs; they
would not be able to jump over the fences so readily; and he acted upon
that idea。 He killed his old ram; and as soon as the young one arrived
at maturity; he bred altogether from it。 The result was even more
striking than in the human experiment which I mentioned just now。
Colonel Humphreys testifies that it always happened that the offspring
were either pure Ancons or pure ordinary sheep; that in no case was
there any mixing of the Ancons with the others。 In consequence of
this; in the course of a very few years; the farmer was able to get a
very considerable flock of this variety; and a large number of them
were spread throughout Massachusetts。 Most unfortunately; howeverI
suppose it was because they were so commonnobody took enough notice
of them to preserve their skeletons; and although Colonel Humphreys
states that he sent a skeleton to the President of the Royal Society at
the same time that he forwarded his paper; I am afraid that the variety
has entirely disappeared; for a short time after these sheep had become
prevalent in that district; the Merino sheep were introduced; and as
their wool was much more valuable; and as they were a quiet race of
sheep; and showed no tendency to trespass or jump over fences; the
Otter breed of sheep; the wool of which was inferior to that of the
Merino; was gradually allowed to die out。
You see that these facts illustrate perfectly well what may be done if
you take care to breed from stocks that are similar to each other。
After having got a variation; if; by crossing a variation with the
original stock; you multiply that variation; and then take care to keep
that variation distinct from the original stock; and make them breed
together;then you may almost certainly produce a race whose tendency
to continue the variation is exceedingly strong。
This is what is called 〃selection〃; and it is by exactly the same
process as that by which Seth Wright bred his Ancon sheep; that our
breeds of cattle; dogs; and fowls; are obtained。 There are some
possibilities of exception; but still; speaking broadly; I may say that
this is the way in which all our varied races of domestic animals have
arisen; and you must understand that it is not one peculiarity or one
characteristic alone in which animals may vary。 There is not a single
peculiarity or characteristic of any kind; bodily or mental; in which
offspring may not vary to a certain extent from the parent and other
animals。
Among ourselves this is well known。 The simplest physical peculiarity
is mostly reproduced。 I know a case of a man whose wife has the lobe of
one of her ears a little flattened。 An ordinary observer might
scarcely notice it; and yet every one of her children has an
approximation to the same peculiarity to some extent。 If you look at
the other extreme; too; the gravest diseases; such as gout; scrofula;
and consumption; may be handed down with just the same certainty and
persistence as we noticed in the perpetuation of the bandy legs of the
Ancon sheep。
However; these facts are best illustrated in animals; and the extent of
the variation; as is well known; is very remarkable in dogs。 For
example; there are some dogs very much smaller than others; indeed; the
variation is so enormous that probably the smallest dog would be about
the size of the head of the largest; there are very great variations in
the structural forms not only of the skeleton but also in the shape of
the skull; and in the proportions of the face and the disposition of
the teeth。
The Pointer; the Retriever; Bulldog; and the Terrier; differ very
greatly; and yet there is every reason to believe that every one of
these races has arisen from the same source;that all the most
important races have arisen by this selective breeding from accidental
variation。
A still more striking case of what may be done by selective breeding;
and it is a better case; because there is no chance of that partial
infusion of error to which I alluded; has been studied very carefully
by Mr。 Darwin;the case of the domestic pigeons。 I dare say there may
be some among you who may be pigeon 'fanciers'; and I wish you to
understand that in approaching the subject; I would speak with all
humility and hesitation; as I regret to say that I am not a pigeon
fancier。 I know it is a great art and mystery; and a thing upon which a
man must not speak lightly; but I shall endeavour; as far as my
understanding goes; to give you a summary of the published and
unpublished information which I have gained from Mr。 Darwin。
Among the enormous variety;I believe there are somewhere about a
hundred and fifty kinds of pigeons;there are four kinds which may be
selected as representing the extremest divergences of one kind from
another。 Their names are the Carrier; the Pouter; the Fantail; and the
Tumbler。 In the large diagrams they are each represented in their
relative sizes to each other。 This first one is the Carrier; you will
notice this large excrescence on its beak; it has a comparatively small
head; there is a bare space round the eyes; it has a long neck; a very
long beak; very strong legs; large feet; long wings; and so on。 The
second one is the Pouter; a very large bird; with very long legs and
beak。 It is called the Pouter because it is in the habit of causing
its gullet to swell up by inflating it with air。 I should tell you that
all pigeons have a tendency to do this at times; but in the Pouter it
is carried to an enormous extent。 The birds appear to be quite proud
of their power of swelling and puffing themselves out in this way; and
I think it is about as droll a sight as you can well see to look at a
cage full of these pigeons puffing and blowing themselves out in this
ridiculous manner。
The third kind I mentionedthe Fantailis a small bird; with
exceedingly small legs and a very small beak。 It is most curiously
distinguished by the size and extent of its tail; which; instead of
containing twelve feathers; may have many more;say thirty; or even
moreI believe there are some with as many as forty…two。 This bird
has a curious habit of spreading out the feathers of its tail in such a
way that they reach forward; and touch its head; and if this can be
accomplished; I believe it is looked upon as a point of great beauty。
But here is the last great variety;the Tumbler; and of that great
variety; one of the principal kinds; and one most prized; is the
specimen represented herethe short…faced Tumbler。 Its beak is
reduced to a mere nothing。 Just compare the beak of this one and that
of the first one; the CarrierI believe the orthodox comparison of the
head and beak of a thoroughly well…bred Tumbler is to stick an oat into
a cherry; and that will give you the proper re